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In addition, a study showed that 5-azacitidine can significantly increase the tri-methylation of lysine-4 on histone H3 in mouse primary spermatocytes, which indicated that 5-azacitidine could modulate histone methylation (Burlibaa et al., 2021)

DAPI was not used to stain the nuclei since its adsorption to the transwell filter background impairs the visualization of VE-Cad- Alexa Fluor 488 signal