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and R35GM122510 to E.H.E.), the National Research Foundation of Korea (2021R1A6A1A10042944, 2017R1A5A1015366, 2020R1C1C1008823) to S.Y.P., and NSFC/RGC Joint Research Scheme N_CityU104/19 and Hong Kong Research Grant Council Collaborative Research Fund: C6021-19E to J.F

In THP-1 macrophage-derived foam cells, leonurine promoted cholesterol efflux and alleviated cellular lipid accumulation in response to the PPAR/LXR signalling pathway, and it prevented atherosclerosis in mice deficient in apoE/ mice (Jiang et al., 2017)