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It follows that neurons that sense metabolic needs to maintain energy balance 19,20,21 and neuronal populations involved in reward-based feeding 22,23,24,25 , along with non-neuronal cells 26,27,28,29 respond to hormones regulating food intake and energy balance, for instance via nutrient sensing 30

1-Methyl-4-phenylpyridinium (MPP+, Sigma-Aldrich, USA, D048) was dissolved in dimethyl sulfoxide (DMSO, Sigma-Aldrich, USA, D2650) to a final concentration of 0.01%