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doi: 10.1007/s11010-018-3332-x

Liver specification occurs around embryonic day 8.5, triggered by BMP signaling from mesenchymal cells of the septum transversum and FGF from the cardiac mesoderm, under the influence of transcription factors Hepatocyte Nuclear Factor-1 (HNF-1), Forkhead box A1 (FOXA1), FOXA2, and GATA Binding Protein 4 (GATA4) (23, 39, 40)