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In contrast, we observed a higher arginine/proline media ratio (higher metabolite concentrations in arginine medium compared to proline medium) for succinate, cis -aconitate, citrate, norleucine, -alanine, creatinine, citramalate, lysine, aspartate, isoleucine, and 10,13-dimethyltetradecanoate in gdh3 / gdh3 mutants compared to wild type (Fig