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Causes of vitamin B12 deficiency include food cobalamin malabsorption (low intestinal absorption of the vitamin from food) due to intrinsic factor receptor deficiency (a substance needed for absorption of B12), impaired calcium availability, gastrointestinal disorders such as chronic gastritis, Crohns disease, and other bowel inflammatory disorders, surgery on the digestive tract such as total or partial gastrectomy, or autoimmune conditions

Bacterial burden was subsequently quantified by manual colony counting, with results expressed as CFU per organ

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The Morris water maze task, a spatial memory test, involved an 8-day acquisition phase and a Day 9 probe trial