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At the mitochondrial outer membrane, BCL2/adenovirus E1B 19 kDa protein-interacting protein 3 (Bnip3) and NIX interact with microtubule-associated proteins 1A/1B light chain 3 (LC3) of the autophagosome, which not only mediated outer membrane permeability but also acted as a mitophagy initiation signal [88]