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In cereal grains, ROS mediate programmed cell death (PCD) in the aleurone layer through cross talk with GA and ABA ( 2+ signaling ( 2 O 2 also enhances GA-induced -amylase synthesis in aleurone cells, whereas antioxidants such as CAT, APX, and SOD suppress it ( 2 O 2 enhances germination and alleviates ABA-induced inhibition in Arabidopsis and pea seeds through GA-dependent pathways (Singh et al., 2016)